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  • Willmott K, Lamas G, Hall J, Vitale F, Boyer P, Petit J-C, Radford J, Elias M. A new species and thirty-eight new subspecies of equatorial Ithomiini (Lepidoptera, Nymphalidae, Danainae). Tropical Lepidoptera Research. 2021;31:1–80. doi:10.5281/ZENODO.5526837
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  • Willmott K, Marín M, Nakahara S, Pomerantz T, Lamas G, Huertas B, Espeland M, Xiao L, Hall J, Willmott J, et al. A revision of the new Andean butterfly genus Optimandes Marín, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina). Tropical Lepidoptera Research. 2019;29:29–44. doi:10.5281/ZENODO.2650482
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