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Pyrcz T, Stachowicz I, Zubek A, Espeland M, Jiménez O, Lachowska-Cierlik D, Florczyk K. A New Species of Butterfly of the Genus Protopedaliodes from the Isolated Tramen Tepui in the Guyana Shield (Lepidoptera, Nymphalidae, Satyrinae). Neotropical Entomology. 2021;50:218–228. doi:10.1007/s13744-020-00830-6

Padrón PS, Pyrcz T, Willmott K. A Contribution Towards Resolving the Systematics of the High-Altitude Tropical Andean Satyrine Genus Altopedaliodes Forster, 1964 (Lepidoptera, Nymphalinae: Satyrinae). Neotropical Entomology. 2021;50:767-803. https://link.springer.com/10.1007/s13744-021-00897-9. doi:10.1007/s13744-021-00897-9