Pyrcz T, Stachowicz I, Zubek A, Espeland M, Jiménez O, Lachowska-Cierlik D, Florczyk K. A New Species of Butterfly of the Genus Protopedaliodes from the Isolated Tramen Tepui in the Guyana Shield (Lepidoptera, Nymphalidae, Satyrinae). Neotropical Entomology. 2021;50:218–228. doi:10.1007/s13744-020-00830-6
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- Neild A, Losada ME, Willmott K. A distinctive new subspecies of Moschoneura pinthous (Linnaeus, 1758) (Lepidoptera: Pieridae: Dismorphiinae) from the Orinoco Delta, Venezuela, with comments on the species-level taxonomy. 2021. https://zenodo.org/record/4966791. doi:10.5281/ZENODO.496679197
- Bollino M, Racheli L, Racheli T. Parides anchises (Linnaeus, 1758) species-group in Venezuela: distribution and diversification. Atalanta. 2021;52:108-119.76
- Costa M, Viloria ÁL, Callaghan C, Trujano-Ortega M, Neild A, Benmesbah M. Lepidoptera del Pantepui. Parte XI. Nuevos Riodinidae (Riodininae), Pieridae (Dismorphiinae) y Nymphalidae (Satyrinae). Antenor. 2021;8(1):2–28.1
- Padrón PS, Pyrcz T, Willmott K. A Contribution Towards Resolving the Systematics of the High-Altitude Tropical Andean Satyrine Genus Altopedaliodes Forster, 1964 (Lepidoptera, Nymphalinae: Satyrinae). Neotropical Entomology. 2021;50:767-803. https://link.springer.com/10.1007/s13744-021-00897-9. doi:10.1007/s13744-021-00897-996
- González J, Domagała PJ. Castniidae of the Museum of Natural History of the University of Wrocław: new findings from Friedrich Wilhelm Niepelt’s collection with comments on Karl Adolf Georg Lauterbach and August Weberbauer. Nota Lepidopterologica. 2021;44:123-132. https://nl.pensoft.net/article/60261/. doi:10.3897/nl.44.6026171
- Penz CM. Revised species definitions and nomenclature of the blue and purple/rose Cithaerias butterflies (Lepidoptera, Nymphalidae, Satyrinae). Zootaxa. 2021;4963:293-316. https://www.biotaxa.org/Zootaxa/article/view/zootaxa.4963.2.3. doi:10.11646/zootaxa.4963.2.359
- Viloria ÁL. The Hermeuptychia Papers. Anartia. 2021;32:26–52. doi:10.5281/ZENODO.557129223
- Willmott K, Lamas G, Hall J, Vitale F, Boyer P, Petit J-C, Radford J, Elias M. A new species and thirty-eight new subspecies of equatorial Ithomiini (Lepidoptera, Nymphalidae, Danainae). Tropical Lepidoptera Research. 2021;31:1–80. doi:10.5281/ZENODO.552683719
- Rios-Malaver I, Olarte-Quiñonez C, Viloria ÁL. Diversidad de especies y estructura del ensamblaje de mariposas (Lepidoptera: Papilionoidea) en un paisaje de bosque nublado periurbano en la Cordillera de la Costa, Venezuela. Anartia. 2021;31:78–101.24