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  • Costa M, Viloria ÁL, Attal S, Neild A, Fratello S, Benmesbah M. Lepidoptera del Pantepui. Parte VIII Nuevos Nymphalidae (Charaxinae y Satyrinae) y Riodinidae (Riodininae). Anartia. 2019;29:20–48.
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  • Viloria ÁL, Costa M. Butterflies. In: Biodiversity of Pantepui. The Pristine “Lost World” of the Neotropical Guiana Highlands. Academic Press; 2019. p. 34.
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